The earliest flowering plants date back about 130 million years. According to Cronquist Evolution and Classification of Flowering Plants, the most primitive of all living angiosperms belong to the subclass Magnoliidae. This subclass contains several primitive plant families, including the water-lily family (Nymphaeaceae), buttercup family (Ranunculaceae) and Magnoliaceae. Plants in the magnolia family have the following characteristics:

• Large flowers with numerous tepals
• Numerous spirally arranged stamens at the base of a conelike receptacle bearing numerous spirally arranged carpels. At maturity the carpels develop into a woody, conelike aggregate of seed-bearing follicles. Each seed has a fleshy red outer layer (aril) and hangs from its follicle by a threadlike stalk. Other primitive floral characteristics are radial symmetry or actinomorphic (floral parts similar in size & shape), perfect (with functional androecium and gynoecium), complete (with all 4 floral parts: calyx, corolla, androecium & gynoecium), and floral axis (receptacle) elongated. The latter characteristic is clearly visible in the magnolia blossom. With all the woody, spirally arranged carpels (follicles), this axis truly resembles a conelike structure [2].

Star magnolia is one of the hardiest of the genus. It can be trained as a tree growing to 20 feet (as pictured here), or multi-trunked bush with a 15-foot spread.  Typically branching close to the ground, the multi-stemmed form develops with a dense head of foliage. Star magnolia makes a wonderful patio, lawn specimen or accent tree. Lower foliage can be removed to show off the trunk and to create more of a tree form. Otherwise, the persistent lower branches and oval to round form lend a “large bush” look to the plant. The white flowers have a slight touch of pink, and are produced in spring before the leaves appear, even on young plants. Flowers are usually not as sensitive to cold as saucer magnolia, but they can still be injured if cold weather arrives during flowering [6].

Star magnolia is intolerant of root competition or dryness, and plants grow slowly, perhaps one foot per year. Plant in the full sun in a rich, porous and slightly acid soil. It is hard to transplant successfully and in the north should be moved balled and burlapped when actively growing. In USDA hardiness zones 7 and 8, transplant in late winter while the plants are still dormant, or after the growth flush in the spring or plant from containers at any time [6].

Taxonomic relations in the family Magnoliaceae have been puzzling botanists for a long time. Because the family is quite old and has survived many geological events such as ice ages, mountain formation and continental drift, the family's distribution is scattered nearly worldwide. Some species have been isolated for a long time, while others were in close contact and able to exchange gentic material until later in their lineage. To create divisions in the family solely based on morphological characteristics has proven to be an impossible task.

With the advent of DNA sequencing in the late 20th century, intense research into phylogenetic relationships within the family has been ongoing. Long boring explanations of the studies' results can be had at Wikipedia [5].

Magnoliaceae - Magnolia Family
The earliest flowering plants date back about 130 million years. According to Cronquist Evolution and Classification of Flowering Plants, the most primitive of all living angiosperms belong to the subclass Magnoliidae. This subclass contains several primitive plant families, including the water-lily family (Nymphaeaceae), buttercup family (Ranunculaceae) and Magnoliaceae.
Tree Encyclopedia | Tree Index | Magnolia Index