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Tilia americana L. - American Basswood
Tiliaceae -- Basswood family
Commonly called basswood, American Linden. Trees in this family are
commonly known as limes in the UK.
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American basswood (Tilia americana), s a native deciduous
tree. The northernmost Tilia species, it is a large,
rapid-growing tree of eastern and central hardwood
woodlands. Best growth is in the central part of the range
on deep, moist soils; development is vigorous from sprouts
as well as seed. American basswood is an important timber
tree, especially in the Great Lakes States. The soft, light
wood has many uses in wood products. The tree is also well
known as a honey-tree, and the seeds and twigs are eaten by
wildlife. It is commonly planted as a shade tree in urban
areas of the eastern states where it is called American
linden. Mature heights range from 75 to 130 feet (23-40 m)
with diameter ranges from 36 to 48 inches (91-122 cm). The
bark of mature trees is up to 1 inch (2.54 cm) thick at the
base of the trunk. The bark is furrowed into narrow,
flat-topped, firm ridges with characteristic horizontal
cracks; young trees have smooth, thin bark. The
inflorescence is a drooping axillary cyme. The fruit is dry,
hard, indehiscent, subglobose to short-oblong, and is
usually 0.2 to 0.28 inch (5-7 mm) in diameter, and bears one
or two seeds.
Native Range:
American basswood ranges from southwestern New Brunswick and
New England west in Quebec and Ontario to the southeast
corner of Manitoba; south through eastern North Dakota,
South Dakota, Nebraska, and Kansas to northeastern Oklahoma;
east to northern Arkansas, Tennessee, western North
Carolina; and northeast to New Jersey. |
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Cymes and Leafy Bracts (for words like that, you should pay me)
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Climate
Climatic conditions associated with the species range
are generally continental-cold winters, warm summers,
and a humid to subhumid moisture regime. Mean annual
precipitation within the species range is 530 min (21
in) at the western limit and 1140 min (45 in) in the
northeast. The northern limit of basswood approximates
the -
18° to -17° C (0° to 2° F) isotherm for mean daily
minimum January temperature. Basswood reaches its
maximum development in areas averaging 18° to 27° C (65°
to 80° F) in July and receiving 250 to 380 mm (10 to 15
in) of precipitation during the growing season. The
frost-free growing period varies from 80 to 180 days
within its range.
Soils and Topography
Studies relating to the presence of basswood to soil
characteristics in Minnesota, Wisconsin, and Michigan
indicate that stands in which basswood shared dominance
were generally confined to sandy loams, loams, or silt
loams, with basswood obtaining maximum development on
the finer textured soils. Most soils were
classified as Hapludalfs within the Alfisols order,
although some Eutrochrepts (Inceptisols), Cryandepts
(Inceptisols), mesic families of entic Fragiorthods
(Spodosols), and Haplorthods (Spodosols) were noted.
Basswood grows best on mesic sites, but it is also found
on coarse soils such as the sand dunes near Lake
Michigan and on dry, exposed rock ridges in Ontario and
Quebec.
The species grows on soils ranging in pH from 4.5 to
7.5 but occurs more often in the less acidic to slightly
basic part of this range. In fact, calcareous soils have
been associated with the presence of basswood. The
importance of aspect and edaphic factors to local
distribution is reflected by the restriction of basswood
throughout much of its range to moist sites on north-
and east-facing slopes. Maple-basswood forests in
southern Wisconsin are largely restricted to northerly
exposures. Basswood is restricted to more mesic sites in
southern Illinois and in northern Kentucky. At the
western limit of its range, basswood frequently grows on
the eastern side of lakes and along major drainages.
This localized growth is often ascribed to fire
protection. although lack of fire may be a reason for
the persistence of a fire-sensitive species such as
basswood, presence and distribution are controlled more
by soil moisture and the ameliorating effects of water
on the local climate. Basswood is classified as a
nitrogen-demanding species because it grows poorly on
sites deficient in nitrogen. With increasing nitrogen
supplies, basswood growth increases markedly,
approaching a maximum radial increment when 560 to 670
kg/ha (500 to 600 lb/acre) of nitrogen are added.
Basswood leaves have high contents of nitrogen,
calcium, magnesium, and potassium at the time of leaf
fall and they contribute most of these nutrients to the
forest floor. Associated Forest Cover American basswood
grows in mixture with other species and only rarely
forms pure stands. It is dominant in a single forest
type, Sugar Maple-Basswood (Society of American
Foresters Type 26). This cover type is most common in
central Minnesota and western Wisconsin but is
represented elsewhere from central Illinois, northward
to southern Ontario and Quebec, eastward to northwestern
Ohio, and westward along valley slopes of the
prairie forest transition (15). Sugar maple (Acer
saccharum) dominates both overstory and understory
layers, with basswood achieving the position of second
dominant in the tree layer. Common associates are white
ash (Fraxinus americana), northern red oak (Quercus
rubra), eastern hophornbeam (Ostrya virginiana), red
maple (Acer rubrum), and American elm (Ulmus americana). |
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Flowering and Fruiting- The
fragrant, yellow-white, flowers are borne on
loose cymes on long stalks attached to leafy bracts.
Flowering generally occurs in June but can begin in
late May or early July, depending on latitude and
annual variations in temperature. Flowering follows
initial leaf-out and lasts approximately 2 weeks.
During this period, all stages of floral development
are present on a single tree or even in a single
inflorescence (4 to 40 flowers per inflorescence).
The flowers attract a number of insect pollinators.
In a study of the pollination biology, 66 species of
insects from 29 families were identified as
pollinators of the flowers.
Bees and flies were the most
common diurnal pollinators; moths were the primary
nocturnal visitors. The fruit, a nutlike drupe 5 to
10 mm (0.2 to 0.4 in) in diameter, usually contains
one seed but in collections from both open- and
forest-grown trees, 12 percent of the fruit
contained two seeds and less than 1 percent
contained three seeds. The seeds have a
crustaceous seed coat (testa), a fleshy yellowish
endosperm, and a well-developed embryo. A variety of
forms of fruit and seed have been noted, including
egg-shaped, round, onion-shaped, conical, and
pentagonal. Individual trees tend to consistently
produce fruit of a particular form and size.
Seed Production and
Dissemination- Fruits ripen in September and October
and are soon dispersed by such mechanisms as wind,
gravity, and animals. although the flower bracts are
reported to aid in wind dispersal, fruits rarely are
carried more than one or two tree lengths from the
parent. In addition to their limited role in seed
dispersal, bracts may act as "flags" to attract
pollinators (especially nocturnal ones) to the
inflorescences. Animals probably increase the seed
dispersal significantly.
Damaging Agents- Basswood
plantations established on weed infested old-field
sites are susceptible to girdling by mice and voles,
and completely girdled trees die. In a southern
Ontario plantation, 44 percent of the basswood stems
were completely girdled and 39 percent were
partially girdled. The species responsible for
the girdling, the meadow vole, does most of this
damage feeding beneath the snow. Rabbits also feed
heavily on seedlings and small saplings in both
plantations and natural stands.
Basswood seeds are eaten by mice, squirrels, and
chipmunks, thus reducing the chances of seedling
establishment. Many different insects attack
basswood, but few serious insect problems exist. The
linden borer (Saperda vestita) makes long, irregular
tunnels, particularly at the base of the tree, and
may damage weak, very young, or overmature trees.
Local infestations of defoliators may occur. The
primary ones include the linden looper (Erannis
tiliaria), basswood leafminer (Baliosus nervosus),
spring cankerworm (Paleacrita vernata), fall
cankerworm (Alsophila pometaria), whitemarked
tussock moth (Orgyia leucostigma), gypsy moth
(Lymantria dispar), and forest tent caterpillar
(Malacosoma disstria). In New England,
American basswood is a highly preferred host for
gypsy moth,
while in southern Quebec, it was classified as
intermediate in susceptibility to gypsy moth
defoliation. The foliage is
host to various diseases-anthracnose (Gnomonia
tiliae), black mold (Fumago vagans), and leaf spot
(Cercospora microsora)-but none seem to do serious
damage. The wood of basswood decays easily and once
exposed can be host to many of the common hardwood
decay organisms such as the yellow cap
fungi (Pholiota limonella) and Collybia velutipes.
Phellinus igniarius, Ustulina deusta, and nectria
canker (Nectria galligena) also are found on
basswood. Little defect is encountered in basswood
when harvested before it reaches 120 years of age.
Beyond this age, the chances of losses due to decay
are greatly increased. Cull studies in the forests
of Ontario indicate that yellow-brown stringy rot
was the most common bole defect encountered; brown
stain, some incipient yellow rot, and green stain
were also found. The thin bark of this species is
easily damaged by fire. Basswood is one of the
hardwoods least susceptible to late spring frosts.
Special Uses - Basswood has relatively soft wood
that works exceptionally well and is valued for hand
carving and has many other uses including cooperage,
boxes, veneer, excelsior, and pulp. Basswood is
economically important for timber, especially in the
Great Lakes States. The inner bark, or bast, can be
used as a source of fiber for making rope or for
weaving such items as baskets and mats. Basswood
flowers produce an abundance of nectar from which
choice honey is made. In fact, in some parts of its
range basswood is known as the bee-tree. Throughout
the Eastern United States, basswood is frequently
planted along city streets. |
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