Scots Pine or Scotch Pine - Pinus sylvestris
Waterer Scots Pine - Pinus sylvestris ' watereri'
Family: Pinaceae.
Scotch pine is an introduced species in North America, brought here from Europe probably in colonial days. although it is used for both pulpwood and sawlogs, its principal value in the United States appears to be as a Christmas tree, as an ornamental, and for erosion control.
 


This Morton Arboretum specimen shows the bright orange bark of Scots Pine.


These elegant Scots pines are 78 years old, part of the original
Morton Arboretum plantings circa 1924.

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Scotch pine has been widely planted in the United States, especially in the Northeast, Lake States, Central States, and Pacific Northwest. It is now considered naturalized in parts of New England and the Lake States. The species has also been planted across southern Canada.

Scotch pine is the most widely distributed pine in the world. It grows naturally from Scotland almost to the Pacific Ocean and from above the Arctic Circle in Scandinavia to the Mediterranean. Its titleitudinal range is from sea level to about 2440 m (8,000 ft). Scotch pine is adapted to a wide variety of climates as indicated by its extremely large natural range. It grows in areas with an annual precipitation exceeding 1780 mm (70 in) and in areas with an annual precipitation as little as 200 mm (8 in). Scotch pine survives in the Verkhoyansk Mountains of eastern Siberia where winter temperatures have been recorded as low as -83° F. In some areas it grows where the subsoil is permanently frozen. Scotch pine can also survive high temperatures, and it is found at middle titleitudes in the Mediterranean region. The primary distribution of Scotch pine, however, indicates that it is a tree of the continental climates.

Scotch pine, like many of the hard pines, is intolerant of shade. Seedlings germinating under a dense forest canopy do not survive for long. although the seedlings will grow very well on fertile soil, they are usually found on the more sandy dry soils because of the lack of competition from other trees and plants.

During the last century, Scottish foresters have had serious difficulties establishing Scotch pine regeneration under mature pine stands. This difficulty appears to be partly due to grazing by deer and domestic animals. Successful regeneration has been achieved, however, with the uniform or shelterwood compartment system, which also appears to be successful in the Scandinavian countries.

The best regeneration is found in stands with the following characteristics: large seed supply, open or light tree canopy, light understory ground cover, and exposed mineral soil or no continuous layer of raw humus.

In the United States in the Northeastern and Lake States, Scotch pine reproduction is extremely abundant on the more sandy sites. As soon as the parent stand reaches seed bearing age, it begins to spread outward into firebreaks and along open roadsides. In many areas reproduction is so plentiful as to present a mat of seedlings, and this aggressive reproductive habit has concerned foresters who preferred to grow other conifer species on these sites. A recent event in New York may change this reproductive potential of Scotch pine. When scleroderris canker (Gremmeniella abietina) is present in the Scotch pine overstory, the advance reproduction can be completely eliminated. As this disease has advanced across northern New York, the "mats" of Scotch pine reproduction have become seas of dead seedlings. New seedlings continue to germinate under the parent stand but become infected and die within 1 or 2 years. Scleroderris canker is present in Scotland but it is not known whether this disease is related to the reproduction problems there. (1)
 


Scots Pine Cones and Foliage

Flowering and Fruiting- although Scotch pine is primarily a monoecious species, some shoots, branches, and even entire trees are predominantly of one sex. Male flower primordia are formed in late summer at the base of the bud that will make the next year's growth. During the winter their presence can be noted as a slight swelling, and the preferred male catkins are easily visible if a bud is dissected. About 2 weeks after growth begins in the spring, the male catkins enlarge to 0.6 to 0.7 cm. (0.2 to 0.3 in) long and shed pollen. At this time they are yellow.

The male catkins are borne at the base of the twigs, replacing leaf clusters. They are most common in the lower part of the crown and on short lateral twigs. Because they replace leaves, an excess of pollen production can lead to sparse foliage. A Pennsylvania breeder who selected for early flower production for two generations obtained a variety that produced plentiful pollen but few needles and it was worthless as a Christmas tree.

Female flower primordia are also formed in late summer but are microscopic. They are borne at the tips of buds for the next year's growth. There may be one, two, or three on a single bud. They first become visible after the buds expand in the spring. The primordia enlarge into female flowers or strobili about 2 weeks after growth begins in the spring, at a time when the new growth has completed 75 percent of its elongation for the season. Because of this, shearing of the outside branches such as is practiced by Christmas tree growers removes all female flowers. Indeed, trees sheared in June will not produce seed for the next 3.5 years.

Flowering occurs in late May or early June. On any one tree nearly all pollen is shed and nearly all the female flowers are receptive during the same 2- or 3-day period. In any one stand most trees flower within a day or two of each other. Trees of different provenances may differ in blooming time by several days, however; trees of northern provenances bloom the earliest.

Pollen production tends to be concentrated on short lateral twigs in the lower half of a tree crown. Female flowers are borne on the most vigorous shoots. They tend to be concentrated on upper branches but may occur in any part of the crown receiving full sunlight.

Pollination occurs in early summer, at a time when the female strobili are from 0.6 to 0.7 cm (0.2 to 0.3 in) long. Shortly after pollination, the scales of the female strobili thicken, and the pollen grains germinate and send out a short pollen tube. At this time the female strobili become reflexed instead of pointing forward. For the next 12 months the germinated pollen remains dormant and the female strobili grow little. A little more than a year after pollination, the germinated pollen grains renew growth and fertilize the ovules. In June, soon after fertilization, the conelets rapidly elongate and reach full size by early summer. Seeds mature and cones ripen in early October. The cones require titleernating periods of dry and wet weather to open and shed few seed until early winter. Indeed, many seeds are retained on the tree until early spring.

Seeds from any one tree can be sorted visually by color into those that are full and those that are empty-empty seeds are much lighter in color (often nearly white) than full ones. On any one tree the full seeds are fairly uniform in color and size, but both traits vary considerably from tree to tree. Trees from the same stand may produce seeds ranging from tan to almost black and from all one color to speckled. Seed size varies in a geographic pattern-seeds from the extreme northern latitudes are half the size of those from the southern part of the range. (1)


Scots Pine Bark (lower trunk)


 

Damaging Agents- Scotch pine in North America is subject to a number of agents that can severely damage or kill the trees. Some of these agents are not present in Europe and Asia and, as a result, the species has not yet had an opportunity to develop genetic resistance.

Fire and wind can damage the trees. Young stands have thin bark and are heavily damaged by fire. Older trees with thicker bark are moderately resistant. Scotch pine has more branches per whorl than red or white pine and this large number of branches makes the tree weak at the nodes. During severe wind storms, trees may snap off at the nodes 3 to 6 m (10 to 20 ft) above the ground. Wildlife and insects are also damaging. The pine grosbeak feeds on the terminal and lateral buds of Scotch pine causing numerous small crooks. Trees of Scandinavian provenances are heavily attacked. In Christmas tree plantations, this feeding can cause major economic losses; a single year's feeding can reduce the tree harvest by 50 percent. This is a minor problem to timber growers, however. On occasion, porcupine seriously damage Scotch pine plantations by girdling young trees, causing dead tops.

The pine root collar weevil (Hylobius radicis) is a major cause of tree death in young plantations in the Lake States. The weevil girdles the tree at the base, killing it within 3 to 4 years. The damage is especially severe on dry sandy soils. The fast-growing central European trees are particularly susceptible (26). In Michigan, on low quality sites, mortality frequently reaches 70 to 80 percent. The pine root tip weevil (Hylobius rhizophagus) causes serious damage in Michigan on Scotch pine Christmas trees grown from stump culture. These trees result from leaving the lower limbs on cut trees to grow into a second tree crop. The pine root tip weevil larvae feed on the roots and root tips, resulting in reduced height growth and flagged shoots, and eventual death. In some cases the pine root tip weevil and the pine root collar weevil attack some Scotch pine stands simultaneously, causing more mortality than expected from either insect alone.

The European pine sawfly (Neodiprion sertifer) causes moderate damage in Christmas trees and ornamental plantings. Heavy defoliation reduces growth from 10 to 20 percent. The fast-growing Scotch pine variety uralensis shows some resistance to this insect while the slow-growing variety iberica is most susceptible.

Common names:

Scotch pine is also a host for brown spot needle disease of southern pines (Scirrhia acicola). This disease, like Lophodermium, causes premature defoliation and is primarily limited to Christmas tree plantations. The long needle provenances are also more resistant to this disease.

Western gall rust (Endocronartium harknessii) is common on Scotch pine in the Lake States and the Northeast. Individual trees may have several hundred galls. In most cases damage is limited to branch mortality and growth loss. As described earlier, Scotch pine is susceptible to scleroderris canker. This disease is present in many areas in Europe, and as a result, certain Scotch pine provenances show some resistance. Scotch pine is more resistant to scleroderris canker than red pine, and in some areas, red pines have been eliminated from the stand while Scotch pines are still alive. Scleroderris canker can be spread on cut Scotch pine Christmas trees. Therefore, State quarantines have been established to prevent the movement of this disease into noninfected areas.

When southern seed sources of Scotch pine are planted too far north of their normal range, severe foliage winter injury develops. This winter injury causes both branch and tree mortality. In the Lake States, a large number of Christmas tree plantations have been destroyed by this problem. Many of these problems in Scotch pine plantations are the result of planting this species on very poor sites or planting the wrong seed source. Scotch pine has the inherent ability to produce excellent, straight-boled stands under the proper conditions. Hundreds of Scotch pine plantations throughout the Lake States and the Northeast are equal to or better than the best red pine stands. When Scotch pine is planted on very poor sites, however, or when improper seed sources are used, damage by insects is so severe as to make the final stand useless for timber production. (1)



26-year-old Scotch Pine

Economic importance:

More:

(2)

 

Scots Pine shoot, in spring, with two-year old seed cone
(open, brown), one-year old seed cone (green), and
new seed cones (red) and pollen cones (yellow).
Scotch pine is the most widely planted pine introduced in North America. It is also the preferred large-volume Christmas tree in the United States- approximately 30 percent of the 35 million Christmas trees harvested annually are Scotch pine (20).

Because it survives on poor droughty sites, Scotch pine has been used to control erosion in many areas. However, the poor vigor of many of these stands on dry, infertile sites has made them susceptible to serious insect attack and many of them have little potential to produce timber (28).

Scotch pine has also been used to a large extent in ornamental plantings. It grows better than red pine on compacted clay soils frequently found around home sites. Because Christmas tree plantations are a ready source of trees, many trees are removed from these plantations as ornamental stock. Many Scotch pine have also been planted along roadsides throughout the Lake States. Scotch pine is similar in fiber and wood characteristics to red pine and is usable for both pulpwood and saw logs. (1)

 
Morton Arboretum accession 661-65*2 Waterer Scots Pine - Pinus sylvestris 'watereri'

References:
1. USDA Forest Service Pinus sylvestris - Scotch Pine
2. USDA, ARS, National Genetic Resources Program.
Germplasm Resources Information Network - (GRIN) [Online Database]

 

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